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The spheres are ruled by the insect-like Kromon who covet the TARDIS. When Charley is captured she is forced to metamorphosise into a hybrid-insect Queen and so to save her, the Doctor must barter his knowledge of space-travel technology, all the while knowing that he risks opening up all the realms of space to a rapacious race whose creed is not to create, only to plunder.
This story takes place after the TV movie.